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          The highway through the rain forest was completed in 1990, _____ some animal species have died off. 

          A. by when 
          B. since then 
          C. since when 
          D. by which time 
          			
          

			
          
				
          
		
		
	
          
		
          
			
          
				
          
				C				
          
			
          
			
          
				
          
			
          
			
          
				
          
			
          
			
          
				
          
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